Luminopteris Ignis, named from Latin for light, wing, and fire, is a bioluminescent cave-dwelling butterfly with delicate, translucent wings up to 10 cm tall, featuring an intricate venation network. Its wings show glowing orange phosphorescent markings along outer margins contrasting with transparent central regions with faint blue-gray bands. The lightweight body (0-2 g) is adapted for fluttering in temperate caves (20-30ºC). Bioluminescence emits a warm orange-yellow glow (570-620 nm) on wing edges and body, likely for communication and mate attraction. It belongs to the bioluminescent Lepidoptera within the rare Phosphorescent Morphidae cluster, highlighting evolutionary adaptations to subterranean life and expanding knowledge of lepidopteran biodiversity and biofluorescence.