Photonica Luminaris, a bioluminescent butterfly with a 0-10 cm wingspan and body mass up to 10 g, features translucent dark wings adorned with electric blue and fiery orange luminescent patterns along lower edges and veins. Its scalloped hindwings display intricate vein networks with faint glowing traces; antennae tips emit subtle orange light aiding intraspecific communication. Fluorescing at 470-530 nm in the blue-green spectrum, it navigates and signals within deep aphotic cave systems. Preferring stable 10-20°C environments, it employs sustained hovering flight in narrow chambers. Classified in Lepidoptera, with 10 related rare, protected species adapted to subterranean habitats, its bioluminescence likely evolved for predator deterrence and mating signals, highlighting its specialized niche in bioluminescent lepidopteran fauna.